Bearded seals in the Atlantic Arctic: review of post 2010 knowledge available for informing stock assessments


  • Nicolai Scherdin North Atlantic Marine Mammal Commission
  • Jana Djukarić North Atlantic Marine Mammal Commission
  • Geneviève Desportes North Atlantic Marine Mammal Commission



Bearded seal, Erignathus barbatus, Atlantic Arctic, post-2010 knowledge, Literature review


The last extensive (pan-Arctic) review on knowledge available on the bearded seal (Erignathus barbatus) was conducted by Cameron et al. in 2010. As bearded seals are hunted off Svalbard and Greenland but no stock assessments are available, NAMMCO requested a status review, and if possible, an assessment of the species in its area. This literature review attempts to summarise the knowledge that has become available post 2010, with a focus on the Atlantic Arctic. A large amount of information has become available on the behaviour of the bearded seal, with hearing, vocalisation, haul-out behaviour and movement patterns (through satelitte tagging), and their phenology, being well studied. A database of baseline blood parameters is slowly being built but is still limited. New data on distribution has emerged from PAM studies and non-targeted surveys. Abundance estimates are missing for Svalbard, but partial estimates have become available for the North Water Polynya in 2009 and 2014. Additionally, observations of bearded seals from aerial line-transect surveys are available for several areas of Greenland but have not been analysed. More information has become available on the impact of anthropogenic stressors, such as climate change and other related environmental changes, although demographic impacts of changes are missing. Catch data exists for both Svalbard and Greenland, but for the latter the data needs to be thoroughly validated. In summary, information on stock structure as well as local and global abundance estimates, which are both important to assess the sustainability of current catches, are still missing. However other lines of evidence can inform the delineation of management areas and the results of a pan-arctic genetic study should become available shortly. There is survey data available from Greenland that could be used to generate local abundance estimates, the analysis of which should be prioritised.


Agafonov, A. V., & Chernetsky, A. D. (2018). Underwate Acoustic Signaling of Bearded Seals (Erignathus barbatus) in the White Sea. Oceanology, 58(6), 847–855.

Andersen, M., Hjelset, A. M., Gjertz, I., Lydersen, C., & Gulliksen, B. (1999). Growth, age at sexual maturity and condition in bearded seals (Erignathus barbatus) from Svalbard, Norway. Polar Biology, 21(3),179–185.

Artsdatabanken (2015). Erignathus barbatus (Erxleben, 1777).

Bengtsson, O., Hamilton, C. D., Lydersen, C., Andersen, M., & Kovacs, K. M. (2021). Distribution and habitat characteristics of pinnipeds and polar bears in the Sval-bard Archipelago, 2005-2018. Polar Research, 40.

Boertmann, D., & Bay, C. (2018). Grønlands Rødliste 2018 – Fortegnelse over grønlandske dyr og planters trusselstatus (Greenland's Red List 2018 - List of the threatened status of Greenlandic animals and plants). Aarhus University, DCE – Danish Centre for Environment and Energy and Greenland Institute of Natural Re-sources.

Boertmann, D., & Mosbech, A. (eds) (2017) Baffin Bay. An updated strategic Environmental Impact Assessment of petroleum activities in the Greenland part of Baffin Bay. Scientific report from DCE - Danish Centre for Envi-ronment and Energy No. 218. Aarhus University, DCE – Danish Centre for Environment and Energy.

Boertmann, D., & Rosing-Asvid, A. (2014). Seabirds and seals in Southeast Greenland. Results from a survey in July 2014. Scientific Report from DCE – Danish Centre for Environment and Energy No. 117. Aarhus University, DCE – Danish Centre for Environment and Energy.

Born, E. W., Teilmann, J., & Riget, F. (2002). Haul-out activity of ringed seals (Phoca hispida) determined from satellite telemetry. Marine Mammal Science, 18(1), 167–181.

Boye, T. K., Simon, M. J., Laidre, K. L., Rigét, F., & Staf-ford, K. M. (2020). Seasonal detections of bearded seal (Erignathus barbatus) vocalizations in Baffin Bay and Davis Strait in relation to sea ice concentration. Polar Biology, 43(10), 1493–1502.

Breed, G. A., Cameron, M. F., Ver Hoef, J. M., Boveng, P. L.., Whiting, A., & Frost, K. J. (2018). Seasonal sea ice dynamics drive movement and migration of juvenile bearded seals Erignathus barbatus. Marine Ecology Progress Series, 600, 223–237.

Burns, J. J., & Frost, K. J. (1979). The natural history and ecology of the bearded seal, Erignathus barbatus. Final Report Outer Continental Shelf Environmental Assess-ment Program. Alaska Department of Fish and Game. 311-392.

Cameron, M. F., Bengtson, J. L., Boveng, P. L., Jansen, J. K., Kelly, B. P., Dahle, S. & Wilder, J. M. (2010). Status review of the bearded seal (Erignathus barbatus). U.S. Dep. Commer., NOAA Tech. Memo. NMFS-AFSC-211, 246 p.

Cameron, M. F., Frost, K. J., Ver Hoef, J. M., Breed, G. A., Whiting, A. V., Goodwin, J., & Boveng, P. L. (2018). Habi-tat selection and seasonal movements of young bearded seals (Erignathus barbatus) in the Bering Sea. PLoS One, 13(2), Article e0192743.

Chambellant, M., Lunn, N. J., & Ferguson, S. H. (2012). Temporal variation in distribution and density of ice-obligated seals in western Hudson Bay, Canada. Polar Biology, 35(7), 1105–1117.

Charrier, I., Mathevon, N., & Aubin, T. (2013). Bearded seal males perceive geographic variation in their trills. Behavioral Ecology and Sociobiology, 67(10), 1679–1689.

Cleator, H.J. (1996). The status of the bearded seal, Erignathus barbatus, in Canada. Canadian Field Natural-ist, 110(3), 501–510.

Conn, P. B., T., Moreland, E. E., London, J. M., Cameron, M. F., & Boveng, P. L. (2014). Estimating multispecies abundance using automated detection systems: ice-associated seals in the Bering Sea. Methods in Ecology and Evolution, 5(12), 1280–1293.

Correa, L., Castellini, J. M., Quakenbush, L. T., & O'Hara, T. M. (2015). Mercury and selenium concentrations in skeletal muscle, liver, and regions of the heart and kidney in bearded seals from Alaska, USA. Environmental toxi-cology and chemistry, 34(10), 2403–2408.

COSEWIC - The Committee on the Status of Endangered Wildlife in Canada (2007). COSEWIC Annual Report 2006–2007.

Crain, D.D., Karpovich, S.A., Quakenbush, L., & Polasek, L. (2021). Using claws to compare reproduction, stress and diet of female bearded and ringed seals in the Bering and Chukchi seas, Alaska, between 1953–1968 and 1998–2014. Conservation Physi-ology, 9(1).

Davis, C. S., Stirling, I., Strobeck, C., & Coltman, D. W. (2008). Population structure of ice- breeding seals. Mo-lecular Ecology, 17(13), 3078–3094.

de Vincenzi, G., Parisi, I., Torri, M., Papale, E., Mazzola, S., Nuth, C., & Buscaino, G. (2019). Influence of environ-mental parameters on the use and spatiotemporal distri-bution of the vocalizations of bearded seals (Erignathus barbatus) in Kongsfjorden, Spitsbergen. Polar Biology, 42(7), 1241–1254.

De Vreese, S., van der Schaar, M., Weissenberger, J., Erbs, F., Kosecka, M., Sole, M., & Andre, M. (2018). Marine mammal acoustic detections in the Greenland and Barents Sea, 2013-2014 seasons. Scientific Reports, 8.

Eldegard K., Syvertsen, P. O., Bjørge, A., Kovacs, K. M., Støen, O.-G., & van der Kooij, J. (2021, november 24). Pattedyr: Vurdering av storkobbe Erignathus barbatus for Norge (Mammals: Assessment of bearded seal Erig-nathus barbatus for Norway). Norsk rødliste for arter 2021 (Norwegian red list for species 2021). Artsdata-banken.

Erokhina, I., & Kavtsevich, N. (2019). Blood plasma chemistry in White Sea bearded seals across different age groups. Arctic Environmental Research, 19(4), 159–165.

Finley, K. J., & Renaud, W. E. (1980). Marine mammals inhabitating the Baffin Bay north water in winter. Arctic 33(4), 724–738.

Foster, G., Nymo, I. H., Kovacs, K. M., Beckmen, K. B., Brownlow, A. C., Baily, J. L., ... & Whatmore, A. M. (2018). First isolation of Brucella pinnipedialis and detec-tion of Brucella antibodies from bearded seals Erignathus barbatus. Diseases Of Aquatic Organisms, 128(1), 13-20.

Fournet, M. E. H., Silvestri, M., Clark, C. W., Klinck, H., & Rice, A. N. (2021). Limited vocal compensation for ele-vated ambient noise in bearded seals: implications for an industrializing Arctic Ocean. Proceedings of the Royal Society B, 288(1945).

Frouin-Mouy, H., Kowarski, K., Martin, B., & Bröker, K. (2017). Seasonal trends in acoustic detection of marine mammals in Baffin bay and Melville bay, Northwest Greenland. Arctic, 70(1), 59–76.

Gulland, E.M.D., Haulena, M., Fauquier, D., Lander, M.E., Zabka, T., Duerr, R. & Langlois, G. (2002). Domoic acid toxicity in Californian sea lions (Zalophus californianus): clinical signs, treatment and survival. Veterinary Record, 150, 475-480.

Goertz, C. E. C., Reichmuth, C., Thometz, N. M., Ziel, H., & Boveng, P. (2019). Comparative Health Assessments of Alaskan Ice Seals. Frontiers in Veterinary Science, 6(FEB), 4.

Hamilton, C. D., Kovacs, K. M., & Lydersen, C. (2018). Individual variability in diving, movement and activity patterns of adult bearded seals in Svalbard, Norway. Scientific Reports, 8(1), 1–17.

Hamilton, C. D., Kovacs, K. M., & Lydersen, C. Sympatric seals use different habitats in an Arctic glacial fjord. Marine Ecology Progress Series, 615, 205–220.

Hamilton, C. D., Lydersen, C., Aars, J., Acquarone, M., Atwood, T., Baylis, A., … Kovacs, K. M. (2022). Marine mammal hotspots across the circumpolar Arctic. Diversi-ty and Distributions, 28, 2729–2753.

Hamilton, C D, Lydersen, C., Aars, J., Biuw, M., Boltunov, A. N., Born, E. W., … Kovacs, K. M. (2021). Marine mam-mal hotspots in the Greenland and Barents Seas. Marine Ecology Progress Series, 659, 3–28.

Hamilton, C. D., Lydersen, C., Fedak, M. A., Freitas, C., Hindell, M. A., & Kovacs, K. M. (2019). Behavioural on-togeny of bearded seals Erignathus barbatus through the first year of life. Marine Ecology Progress Series, 627, 179–194.

Haver, S. M., Gedamke, J., Hatch, L. T., Dziak, R. P., van Parijs, S., McKenna, M. F., … Klinck, H. (2018). Monitoring long-term soundscape trends in U.S. Waters: The NO-AA/NPS Ocean Noise Reference Station Network. Marine Policy, 90, 6–13.

Heide-Jørgensen, M. P., Burt, L. M., Hansen, R. G., Niel-sen, N. H., Rasmussen, M., Fossette, S., & Stern, H. (2013). The significance of the North Water Polynya to Arctic top predators. Ambio, 42(5), 596–610.

Heide-Jørgensen, M. P., Sinding, MH. S., Nielsen, N. H., Rosing-Asvid, A., & Hansen, R. G. (2016). Large numbers of marine mammals winter in the North Water polynya. Polar Biology, 39(9), 1605–1614.

Helm, R. C., Costa, D. P., DeBruyn, T. D., O’Shea, T. J., Wells, R. S., & Williams, T. M. (2015). Overview of Effects of Oil Spills on Marine Mammals. Handbook of Oil Spill Science and Technology, 455–475.

Hindell, M. A., Lydersen, C., Hop, H., & Kovacs, K. M. (2012). Pre-partum diet of adult female bearded seals in years of contrasting ice conditions. PLoS ONE, 7(5), Article e38307.

Iversen, M., Aars, J., Haug, T., Alsos, I. G., Lydersen, C., Bachmann, L., & Kovacs, K. M. (2013). The diet of polar bears (Ursus maritimus) from Svalbard, Norway, inferred from scat analysis. Polar Biology, 36(4), 561–571.

Johnson, M. L., Fiscus, C. H., Ostenson, B. T., & Barbour, M. L. (1966). Marine mammals. In N. J. Wilimovsky & J. N. Wolfe (Eds.), Environment of the Cape Thompson region, Alaska. (pp. 877–924). U. S. Atomic Energy Com-mission.

Karpovich, S. A., Horstmann, L. A., & Polasek, L. K. (2020). Validation of a novel method to create temporal records of hormone concentrations from the claws of ringed and bearded seals. Conservation Physiology, 8(1), 1–11.

Kienle, S. S., & Berta, A. (2016). The better to eat you with: the comparative feeding morphology of phocid seals (Pinnipedia, Phocidae). Journal of Anatomy, 228(3), 396–413.

Kienle, S. S., Hermann-Sorensen, H., Costa, D. P., Reichmuth, C., & Mehta, R. S. (2018). Comparative feed-ing strategies and kinematics in phocid seals: suction without specialized skull morphology. Experimental Biology, 221(15).

Kovacs, K. M. (2016). Erignathus barbatus. The IUCN Red List of Threatened Species 2016: e.T8010A45225428. Accessed on 04 January 2022.

Kovacs, K. M. (2018). Bearded Seal: Erignathus barbatus. In B. Würsig, J. G. M. Thewissen, & K. M. Kovacs (Eds.), Encyclopedia of Marine Mammals (Third Edition, pp. 83–86).

Kovacs, K. M., Aguilar, A., Aurioles, D., Burkanov, V., Campagna, C., & Gales, N. (2012). Global threats to pinnipeds. Marine Mammal Science, 28(2), 414–436.

Kovacs, K. M., Belikov, S., Boveng, P., Desportes, G., Ferguson, S., & Hansen, R., (2021). State of the Arctic Marine Biodiversity Report (SAMBR) Update: 2021 State of the Arctic Marine Biodiversity Report (SAMBR) Up-date: Marine Mammals. Technical Report. Conservation of Arctic Flora and Fauna International Secretariat: Aku-rey-ri, Iceland.

Kovacs, K. M., Krafft, B. A., & Lydersen, C. (2020). Beard-ed seal (Erignathus barbatus) birth mass and pup growth in periods with contrasting ice conditions in Svalbard, Norway. Marine Mammal Science, 36, 276–284.

Kovacs, K.M., Lydersen, C. & Gjertz, I. (1996). Birth-site characteristics and prenatal molting in bearded seals (Erignathus barbatus). Journal of Mammalogy, 77(4), 1085–1091.

Kovacs, K. M., Lydersen, C., Overland, J. E., & Moore, S. E. (2011). Impacts of changing sea-ice conditions on Arctic marine mammals. Marine Biodiversity, 41(1), 181–194.

Laidre, K., Heide-Jørgensen, M. P., Stern, H., & Richard, P. (2012). Unusual narwhal sea ice entrapments and delayed autumn freeze-up trends. Polar Biology, 35(1), 149–154.

Ladegaard, M., Macauley, J., Simon, M., Laidre, K. L., Mitseva, A., Videsen, S., … Madsen, P. T. (2021). Sound-scape and ambient noise levels of the Arctic waters around Greenland. Scientific Reports, 11(1), 1–15.

Laidre, K.L., Stern, H., Kovacs, K.M., Lowry, L., Moore, S.E., Regehr, E.V., & Ugarte, F. (2015). Arctic marine mammal population status, sea ice habitat loss, and conservation recommendations for the 21st century. Conservation Biology, 29(3), 724-737.

Leblanc, G., Francis, C. M., Soffer, R., Kalacska, M., & de Gea, J. (2016). Spectral Reflectance of Polar Bear and Other Large Arctic Mammal Pelts; Potential Applications to Remote Sensing Surveys. Remote Sensing, 8(4). 273.

Lefebvre, K. A., Quakenbush, L., Frame, E., Huntington, K. B., Sheffield, G., Stimmelmayr, R., … Gill, V. (2016). Prevalence of algal toxins in Alaskan marine mammals foraging in a changing arctic and subarctic environment. Harmful Algae, 55, 13–24.

Llobet, S. M., Ahonen, H., Lydersen, C., Berge, J., Ims, R., & Kovacs, K.M. (2021). Bearded seal vocalisations across seasons and habitat types in Svalbard, Norway. Polar Biology, 44, 1273–1287.

Lomac-MacNair, K., Andrade, J.P., & Esteves, E. (2019). Seal and polar bear behavioral response to an icebreaker vessel in northwest Greenland. Human–Wildlife Interac-tions, 13(2), 277-289.

Lomac-MacNair K., Andrade J. P., & Esteves E. (01–05 March 2021). Seal and polar bear behavioral response to an icebreaker vessel in Northwest Greenland. [Confer-ence presentation). XI International Conference Marine mammals of the Holarctic, online.

Lomac-MacNair, K., Jakobsson, M., Mix, A., Freire, F., Hogan, K., Mayer, L., & Smultea, M. A. (2018). Seal oc-currence and habitat use during summer in Petermann Fjord, northwestern Greenland. Arctic, 71(3), 334-348.

London, J. M., Conn, P. B., Hardy, S. K., Richmond, E. L., ver Hoef, J. M., Cameron, M. F., Boveng, P. L. (2022). Haul-out behavior and aerial survey detectability of seals in the Bering and Chukchi seas.

Lydersen, C., Assmy, P., Falk-Petersen, S., Kohler, J., Kovacs, K.M., & Reigstad, M. (2014). The importance of tidewater glaciers for marine mammals and seabirds in Svalbard, Norway, Journal of Marine Systems, 129, 452-471.

Lydersen, C., Hammill, M.O., & Kovacs, K.M. (1994). Diving activity in nursing bearded seal (Erignathus barba-tus) pups. Canadian Journal of Zoology, 72(1), 96-103.

MacKenzie, K.M., Lydersen, C., Haug, T., Routti, H., Aars, J., & Andvik, C.M. (2022). Niches of marine mam-mals in the European Arctic. Ecological Indicators, 136, 108661.

MacIntyre, K. Q., Stafford, K. M., Conn, P. B., Laidre, K. L., & Boveng, P. L. (2015). The relationship between sea ice concentration and the spatio-temporal distribution of vocalizing bearded seals (Erignathus barbatus) in the Bering, Chukchi, and Beaufort Seas from 2008 to 2011. Progress in Oceanography, 136, 241–249.

Madan, M. M., Latha, G., Ashokan, M., Raguraman, G., & Thirunavukkarasu, A. (2020). Identification of sound-scape components and their temporal patterns in Kongs-fjorden, Svalbard Archipelago. Polar Science, 26, 1–8.

Manning, T. H. (1974). Variations in the skull of the bearded seal. Technical Report. Biological papers of the University of Alaska. 16.

Marcoux, M., Ferguson, S. H., Roy, N., Bedard, J. M., & Simard, M. (2017). Seasonal marine mammal occurrence detected from passive acoustic monitoring in Scott Inlet, Nunavut, Canada. Polar Biology, 40, 1127–1138.

Martien, K. K., Lang, A. R., Taylor, B. L., Rosel, P. E., Simmons, S. E., Oleson, E. M., ... Hanson, M. B. (2019). The DIP delineation handbook: a guide to using multiple lines of evidence to delineate demographically independ-ent populations of marine mammals. (NOAA Technical Memorandum NMFS-SWFSC-622). U.S. Department of Commerce.

Mattmüller, R. M., Thomisch, K., van Opzeeland, I., Laidre, K. L., & Simon, M. (2022). Passive acoustic moni-toring reveals year-round marine mammal community composition off Tasiilaq, Southeast Greenland. The Journal of the Acoustical Society of America, 151(2), 1380–1392.

McHuron, E. A., Williams, T., Costa, D. P., & Reichmuth, C. (2020). Contrasting whisker growth dynamics within the phocid lineage. Marine Ecology Progress Series, 634, 231–236.

Minzyuk, T. V., Kavtsevich, N. N., & Svetochev, V. N. (2015). New data on the blood cell composition of bearded seal. Doklady Biological Scienc-es, 462(1), 152–154.

Moore, S. E., Stafford, K. M., Melling, H., Berchok, C., Wiig, O., Kovacs, K. M., & Richter-Menge, J. (2012). Comparing marine mammal acoustic habitats in Atlantic and Pacific sectors of the High Arctic: year-long records from Fram Strait and the Chukchi Plateau. Polar Biology, 35(3), 475–480.

NAMMCO-North Atlantic Marine Mammal Commission (2015). Report of the 22nd Meeting of the Scientific Com-mittee.

NAMMCO-North Atlantic Marine Mammal Commission (2016). Report of the 23rd Meeting of the Scientific Com-mittee, 4-7 November 2016.

NAMMCO-North Atlantic Marine Mammal Commission (2019). Report of the 27th Meeting of the Council, 3-4 April 2019, Tórshavn, Faroe Islands.

NAMMCO-North Atlantic Marine Mammal Commission (2020). Report of the 28th Meeting of the Council - Heads of Delegation, 26 March 2020, online.

NAMMCO-North Atlantic Marine Mammal Commission (2021). Report of the Scientific Committee Working Group on By-Catch, 15 October 2021, Video Conference.

NAMMCO-North Atlantic Marine Mammal Commission (n.d.). NAMMCO Catch Database. Retrieved July 26, 2021, from:

Nelson, M. A., Quakenbush, L. T., Taras, B. D., & Ice seal Commitee (2019). Subsistence harvest of ringed, beard-ed, spotted, and ribbon seals in Alaska is sustainable. Endangered Species Research, 40, 1–16.

Olnes, J., Crawford, J., Citta, J. J., Druckenmiller, M. L., von Duyke, A. L., & Quakenbush, L. (2020). Movement, diving, and haul-out behaviors of juvenile bearded seals in the Bering, Chukchi and Beaufort seas, 2014–2018. Polar Biology, 43, 1307–1320.

Parisi, I., Vincenzi, G. de, Torri, M., Papale, E., Mazzola, S., Bonanna, A., & Buscaino, G. (2017). Underwater vocal complexity of Arctic Erignathus barbatus in Kongsfjorden (Svalbard). Journal of the Acoustical Society of America, 142(5), 3104–3115.

Pine, M. K., Hannay, D. E., Insley, S. J., Halliday, W. D., & Juanes, F. (2018). Assessing vessel slowdown for reduc-ing auditory masking for marine mammals and fish of the western Canadian Arctic. Marine Pollution Bulletin, 135, 290–302.

Priest, H., & Husher, P.J. (2004). The Nunavut Wildlife Harvest Study. Final report. Nunavut Wildlife Manage-ment Board. Nunavut, Canada.

Quakenbush, L., Citta, J., & Crawford, J. (2011). Biology of the bearded seal (Erignathus barbatus) in Alaska, 1961-2009. National Marine Fisheries Service.

Quakenbush, L. T., Crawford, J., Nelson, M., & Olnes, J. (2019). Ice Seal Movements and Foraging : Village-based Satellite Tracking and Collection of Traditional Ecological Knowledge Regarding Ringed and Bearded Seals. [OCS Study BOEM 2019-079]. U.S. Dept. of the Interior, Bu-reau of Ocean Energy Management, Alaska Outer Conti-nental Shelf Region, Anchorage, AK.

Risch, D., Clark, C. W., Corkeron, P. J., Elepfandt, A., Kovacs, K. M., Lydersen, & Van Parijs, S. M. (2007). Vo-calizations of male bearded seals, Erignathus barbatus: classification and geographical variation. Animal Behav-iour, 73(5), 747–762.

Rosen, D. A. S., Thometz, N. M., & Reichmuth, C. (2021). Seasonal and Developmental Patterns of Energy Intake and Growth in Alaskan Ice Seals. Aquatic Mammals, 47(6), 559–573.

Rosing-Asvid, A., Dietz, R., Teilmann, J., Olsen, M. T., & Andersen, S. M. (2012). Preliminary report about seals and their sensitivity to oil-exploration in South Green-land. Greenland Institute of Natural Resources. in_South_Greenland-1.pdf

Rosing-Asvid, A., Riget, F., Mikkelsen, L., & Dietz, R. (2015). Seals and their sensitivity to oil exploration in eastern Baffin Bay.Ringed and bearded seal tracking in the Melville Bay area. Aarhus University, DCE – Danish Centre for Environment and Energy and Greenland Insti-tute of Natural Resources.

Roth, E. H., Hildebrand, J. A., Wiggins, S. M., & Ross, D. (2012). Underwater ambient noise on the Chukchi Sea continental slope from 2006–2009. The Journal of the Acoustical Society of America, 131(1), 104.

Scherdin, N. L. P., Olsen, M. T., Heide-Jørgensen, M. P., & Hansen, R. G. (2020). Distribution of bearded seals (Erignathus barbatus) in Greenland. [Unpublished bache-lor's thesis]. University of Copenhagen.

Senoo, H., Imai, K., Mezaki, Y., Miura, M., Morii, M., Fujiwara, M., & Blomhoff, R. (2012). Accumulation of Vitamin A in the Hepatic Stellate Cell of Arctic Top Preda-tors. Anatomical Record-Advances in Integrative Anato-my and Evolutionary Biology, 295(10), 1660–1668.

Sills, J. M., Reichmuth, C., Southall, B. L., Whiting, A., & Goodwin, J. (2020). Auditory biology of bearded seals (Erignathus barbatus). Polar Biology, 43, 1681–1691.

Sills, J. M., Ruscher, B., Nichols, R., Southall, B. L., & Reichmuth, C. (2020). Evaluating temporary threshold shift onset levels for impulsive noise in seals. Journal of the Acoustical Society of America, 148, 2973–2986.

Smith, T. G. (1981). Notes on the bearded seal, Erig-nathus barbatus, in the Canadian Arctic. (Report No. Cat. No. Fs 97-6/1042). Canadian Technical Report of Fisher-ies and Aquatic Sciences.

Stempniewicz, L., Goc, M., Kidawa, D., Urbański, J., Hadwiczak, M., & Zwolicki, A. (2017). Marine birds and mammals foraging in the rapidly deglaciating Arctic fjord - numbers, distribution and habitat preferences, Climate change, 140(3), 533-548.

Summarell, C. C. G., Ingole, S., Fish, F. E., & Marshall, C. D. (2015). Comparative Analysis of the Flexural Stiffness of Pinniped Vibrissae. PLOS ONE, 10(7). Article e0127941.

Terhune, J. M. (2019). The underwater vocal complexity of seals (Phocidae) is not related to their phylogeny. Canadian Journal of Zoology, 97(3), 232–240.

Thometz, N. M., Hermann-Sørensen, H., Russell, B., Rosen, D. A. S., & Reichmuth, C. (2021). Molting strate-gies of Arctic seals drive annual patterns in metabolism. Conservation Physiology, 9, 1–14.

Tryland, M., Lydersen, C., Kovacs, K. M., Rafter, E., & Thoresen, S. I. (2021). Serum biochemistry and haema-tology in wild and captive bearded seals (Erignathus barbatus) from Svalbard, Norway. Acta Veterinaria Scan-dinavica, 63(1), 1–7.

Van Parijs, S. M., Kovacs, K. M., & Lydersen, C. (2001). Spatial and Temporal Distribution of Vocalising Male Bearded Seals: Implications for Male Mating Strategies. Behaviour, 138(7), 905–922.

ver Hoef, J. M., Cameron, M. F., Boveng, P. L., London, J. M., & Moreland, E. E. (2014). A spatial hierarchical model for abundance of three ice-associated seal species in the eastern Bering Sea. Statistical Methodology, 17(C), 46–66.

Watanabe, Y., Lydersen, C., Sato, K., Naito, Y., Miyazaki, N. & Kovacs, K.M. (2009). Diving behavior and swimming style of nursing bearded seal pups. Marine Ecology Pro-gress Series, 380, 287-294.

Wilson, S. C., Crawford, I., Trukhanova, I., Dmitrieva, L., & Goodman, S. J. (2020). Estimating risk to ice-breeding pinnipeds from shipping in Arctic and sub-Arctic seas. Marine Policy, 111.




How to Cite

Scherdin, N., Djukarić, J., & Desportes, G. (2022). Bearded seals in the Atlantic Arctic: review of post 2010 knowledge available for informing stock assessments. NAMMCO Scientific Publications, 12.

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